Nice FMRI study showing that working memory delay activity is primarily in the superficial, feedforward, cortical layers while behavioral response-related activity is primarily in deep, feedback layers.
Layer-dependent activity in human prefrontal cortex during working memory
Emily S. Finn, Laurentius Huber, David C. Jangraw, Peter A. Bandettini
doi: https://doi.org/10.1101/425249
This is very consistent with our recent work:
Bastos, A.M., Loonis, R., Kornblith, S., Lundqvist, M., and Miller, E.K. (2018) Laminar recordings in frontal cortex suggest distinct layers for maintenance and control of working memory. Proceedings of the National Academy of Sciences. View PDF