Kraus et al required rats to run on a treadmill during a working memory task.  They dissociated distance traveled vs time spent running by requiring the rats to run for a fixed distance or a fixed amount of time.  This revealed “time cells” in the hippocampus that reflect  the passage of time.

About the Author

The Miller Lab uses experimental and theoretical approaches to study the neural basis of the high-level cognitive functions that underlie complex goal-directed behavior.